That first OCAU system is similar to what I built for mum and dad. It hasn’t shown any weakness yet. Handles BluRay/1080P easily.
The 2nd OCAU system is similar to a HTPC that I just finished helping a mate build with left over parts, once again, does the job.
The one you want to build is closer spec’ed to my HTPC. IMO, it’s overkill but you will always know that you won’t have to upgrade for a very, very long time.
The biggest thing to get your head around with HTPC’s is that nearly any computer could function as one, even with ancient components. My mate ran a single core AMD in his for around 12 months and it still handled 1080P as it was handled by the GPU. It was a bit sluggish though. Dual cores are a minimum.
The Hauppauge MCE Remotes are great, just plug them in and they work. They are also natively supported by MediaPortal if you decide to use that over WMC.
Get the biggest HDDs you can afford, you’ll fill them quickly and then not have any more space inside the case for additional ones.
Looks like you’ve already decided on this, but if you want dual digital tuners, get a single card with dual tuners. I did the opposite, twin single card tuners. Works the same, but you loose an expansion slot. The Leadtek DTV turners are great value, I have used them exclusively over anything else due to the cost. Hauppauge are probably the best, but you really pay for them. Don’t bother with hybrid tuners, nobody cares about analogue any more.
There’s a lot that I’ve missed (rushing a bit due to needing to study, I can’t resist the forums), but if you want any more opinion on other things that I didn’t answer, just give a yell.